Hi Mike,
Welcome back.
Well, since I wrote above, given that both I and my daughter shoot distant wildlife, when she decided to move up from my old bridge camera to a DSLR, we considered all the options and the benefits of 2x (over my 1.5x) crop factor and not wanting weighty camera and lens led her to buy the Panasonic GH2, a M4/3 format camera for which we bought the 14-140 and 100-300 lenses, so she can shoot at 600mm FFE against my 450mm FFE. No plans to change, so it must be permanent.
I'll stick with my 1.5 cf Nikon, but have dabbled with a Mega-zoom bridge camera (Nikon P510) which goes to 1000mm FFE at f/5.9 as a much cheaper and lighter way to get closer than spending between £1k5 and 7k2 on a long lens for the D5000.
In good light, it definitely produces better results than my Nikon DSLR with 70-300mm (450mm FFE) cropped down.
The downsides are;
I cannot push the iso so hard in poor light (my limits are 400iso max on P510 against 2000iso on D5000)
The AF can be maddeningly slow and I miss so many shots
I cannot crop down so much from full image on the P510 at 1000mm as I can with the D5000.
I am currently considering the Canon SX50 which goes to 1200mm at f/5.9, or the Panasonic FZ200, which only goes to 600mm, but does so at f/2.8!
The latter allowing 2+ stops faster shutter speed or lower iso, but with far less benefit in focal length over my DSLR.
Decisions, decisions
Cheers,

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So to cut a long story short I bought an Olympus OMD-EM5 instead of upgrading the aps 300D. That came with the 12-50mm zoom. I am getting the feeling that this lens isn't as good as the Pen 14-42mm. Not sure yet. It's also a lot longer than the 14-42mm. The EM5 can be bought with that lens. It's very compact but not water proof.
I just assumed crop factor related to area without checking as that would be a sensible way to do it - max print size for instance in a round about way. The correct way must be via pixel size as you imply. The fact that a 16mp camera asks more of a lens than a 12mp one is still factual though.
Also on this shot I am rather surprised how much difference calibration and the increased dynamic range of my new monitor makes to the detail in the blacks. It also shows how much detail pens bury in blacks even in jpg's. From reviews even more is available in the OM.
